Old Growth Forests - Characteristics and Management Implications

Good news. There are still isolated tracts of forest in Maryland which posses both the characteristics of old growth and most of the species that were present when Europeans first came to America. While the majority of these stands are not ³virgin,² which means never touched by axe or saw, they were nonetheless lightly cut and are returning to the condition which ecologists recognize as old growth.

Four Stages of Forest Development

What is ³old-growth² and how does it differ from the woods you generally see? To best explain old growth it is necessary to understand the four basic successional phases of a forest.

Stage 1: Reorganization
The first stage occurs immediately after a disturbance in the forest caused historically by fire and wind or today also by people. This stage is termed reorganization and generally last less than twenty years during which time many nutrient cycles and other soil processes act to stabilize the area. As this stage is completed, seedlings and sprouts of woody plants striving to become the canopy of the new forest shade the soil.

Stage 2: Aggradation
The second stage, aggradation, may last for well over a century as the trees rapidly grow in height, as well as girth. Trees accumulate nutrients and biomass faster during this stage than any other. Thus, to maximize the economic return, nearly every acre of forest commercially managed for wood products is cut during this early stage in succession. Clearcutting of forest in the aggradation stage is the preferred silvicultural method to start the regeneration of a new stand.

Stage 3: Transition
However, if forests are permitted to continue to evolve naturally, the trees will begin to slow their rate of growth as their crowns compete for sunlight and their roots compete for water and nutrients. Shading from neighboring trees, disease, insects, fungi, as well as damage from tree falls, begin to thin out the trees in the forest. Less and less of the nutrients in the ecosystem are tied up in living tissue; nutrients increasingly accumulate in organic matter on and in the forest soil. As the soils enrich and the amount of dead wood accumulates in the system, the abundance and diversity of life responds to the increasing food resources and habitat complexity. This third stage of forest succession is termed transition to denote these changes.

Stage 4: Steady State
Eventually, in less than two hundred years from the initial disturbance, the forest¹s nutrient levels and biomass stabilize so that a steady state, the final stage of forest development, will result. Now the death of a tree is balanced by the germination of a number of seedlings. Over time throughout the stand, the nutrients and life entering the system roughly equal the nutrient loss and death in the forest. A complete life cycle is established and maintained. This condition, called ³old-growth² by ecologists, will exist indefinitely until humans or natural occurrences disturb this balance.

Old-Growth Characteristics

Old-growth forests differ significantly from forest managed for wood products, those maintained to the aggradation stage. As compared to younger forest, old-growth forest are characterized by larger trees, more standing dead trees which are important for wildlife, more downed logs, a more uneven canopy with many gaps and different tree sizes, greater variation in the light level, temperature (mostly cooler), and moisture (more humid) on the forest floor, and greater complexity in overall forest structure (multiple, indistinct growth layers, and greater heterogeneity). Because of these differences and complexities, the diversity and abundance of life supported in an old-growth ecosystem is far greater than that found in the early successional stages of a forest.

Maryland's Heritage

Inability to interpret old growth conditions, priority consideration for retention during the land development process, destruction of old-growth ecosystems and the fragmentation and simplification of the forest is compromising some plant and animal species, including songbirds that require interior forest conditions.

The Maryland Natural Heritage Program and others are beginning to identify and study remaining patches of the State¹s old-growth forest. So far, research has located only a few completely virgin tracts larger than 100 acres. However, they have identified a good number tracts of forest containing old-growth, as well as larger areas (often surrounding pockets of old-growth) where the forest was only cut once or twice, often selectively, (highgrading) and is now in transition to the steady state.

We need to seriously consider the protection of old-growth forest conditions in Maryland. We also need to adequately protect Maryland's biological diversity by setting aside ecologically viable blocks of contiguous forest to allow it to revert to old growth.

Although unintentional, the Maryland Forest Conservation Act (FCA) has inadvertently promoted old growth throughout the State. Since 1991 when enforcement of the Act began, the average forest set aside (not including forested floodplain) of any given development tract has been approximately 20 percent. The language regarding their long-term protection has in general, only recognized passive recreation as a legitimate use. This will result in numerous small tracts being allowed to revert to old growth over ecological time. Less than 5 percent of all dedicated FCA lands include language for lumbering, which is a landowners legal right, if they should choose to do so. This is because a cut forest meets the legal definition of a standing forest according to the Act.

ESA, Inc is heavily involved with performing all aspects of forest stand delineation and forest conservation plans for our clients. We have observed that when a client has land with priority forest retention areas that include floodplain, wetlands, steep slopes and/or highly erodible soils and still are not at desired threshold set-asides or break-even point, that we then consider forest age as possibly the next most important criterion for retention. A 60-year old forest is inherently more valuable than a 30-year old forest. It may take upwards of 60 years + for a stand to reflect an endemic shrub layer. The next most important retention consideration would be to retain forest in a manner that would maximize the total forest area, versus increasing the edge effect.

ESA, Inc. has developed numerous ecological restoration plans for forest retention areas, receiving aff or reforestation credit by improving upon existing conditions, before attempting to create new forest situations. We have been able to manage for nuisance alien plants or taken a sapling stocked (1 to 5-inches diameter) stand and advance seral succession by thinning (crown touching) to push basal growth, thereby creating a more attractive forest setting during the life expectancy of residents who may abut an FCA subdivision greenspace dedication.